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More info about phylum chordata. |
All chordates have a number of structures in common: A notochord (noto = the back; chord = string) is present in all embryos, and may be present or absent/reduced in adults. A dorsal, hollow, ectodermal nerve cord (compare with Annelida and Arthropoda which have ventral, solid, mesodermal nerve cords) typically forms by an infolding of the ectoderm tissue, which then “pinches off” and becomes surrounded by mesoderm. Spinal bifida is the failure of the nervous system to close. The pharangeal slits (pharynx = throat) originally functioned in filter feeding: water is taken into the mouth and let out via the pharangeal slits. The slits filter out food particles and keep them in the animal’s body so they can be put into the digestive tract. In fish, these have become modified as gills, and in humans as our ears and eustachian tubes. A postanal tail (post = behind, after; anal refers to the anus) is present and extends behind the anus in many taxa, thus the anus isn’t at posterior tip of body. In humans, the tail is present during embryonic development, but is subsequently resorbed. Subphylum Cephalochordata (cephalo = head), which doesn’t have a “head.” This subphylum includes Lancets, so-named for their shape. These are in genus Branchiostoma (branchio = gill, fish; stoma = mouth) which was formerly know as genus Amphioxus (amphi = on both sides, double) Lancets look a lot like the generic chordate described above. They have a notochord, but no bones. They are about one inch long, and live in the muddy ocean floor. Subphylum Urochordata (uro = tail) are called the tunicates. Their larvae show typical chordate characteristics, but the adults have lost many of these organs, and what they do still have have become highly-modified. Subphylum Vertebrata may have come from an Amphioxus-like ancestor, however vertebrates have a definite head and Amphioxus doesn’t. Another theory suggests that vertebrates may have come from a larval form similar to tunicate larvae that were capable of reproduction in the larval stage, that is, they are sexually mature prior to metamorphosis, which is referred to as paedogenesis (paedo = child — same root as in pediatrician; genesis = origin, birth), defined as the precocious attainment of sexual maturity in a larva. Vertebrate characteristics include: vertebrae, the skeletal units surrounding the nerve cord a brain, enclosed within a skull an endoskeleton which will grow along with the animal (unlike arthropods which must molt). a closed circulatory system with a ventral heart excretion via kidneys separate males and females with sexual reproduction in most, with a few cases of parthenogenesis. The Classes in Subphylum Vertebrata include: Class Agnatha (a- = not, without; gnatho = jaw) which is the lampreys. They do not have jaws, are eel-shaped, prey on fish, and have larval forms which are different from the adults. |