Tubeless tires have softer and more flexible sidewalls than tube-type tires (except for some very serious low profile autocross tires with steel reinforced sidewalls). Modern radials tend to squat under load and make the sidewalls bulge out a bit, This puts a larger footprint on the road, and the softer sidewall reduces rolling resistance, reduces heating, and improves fuel economy. When you install a tube in a tubeless tire you sort of defeat the advantages of the flexible sidewall, and the car will ride stiffer (more harsh on bumps).
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